De Villiers was appointed captain of South Africa's one-day and Twenty20 teams when Gary Kirsten was named coach last June, but a hand injury kept him out of matches against Australia.
Hashim Amla took over as captain in a drawn Twenty20 series and a one-day series which Australia won 2-1.
It will be a tough baptism for De Villiers, one of South Africa's key batsmen, because he has also been selected to keep wicket.
Sri Lanka, coming off a 2-1 Test series defeat, will seek to recapture the form that took them to the World Cup final last year.
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